The study, published April 29 in the journal Global Ecology and Conservation, also found that three other species were good indicators of a depleted forest state: the mustached babbler (Malacopteron magnirostre), the brown-cheeked bulbul (Alophoixus bres) and the chestnut-backed scimitar babbler (Pomatorhinus montanus). The white-bellied erpornis (Erpornis zantholeuca), meanwhile, was found to be a good indicator species for mixed gardens, while none were indicators for oil palm farms.